Optimal Optimus, a wise and compassionate Maximal commander, developed his trademark valor and heroism during his role in the Beast Wars. Exposure to a nebulous entity reconfigured him into the ultimate four-mode Transmetal. Optimal Optimus' reactive smart missile cannons are functional in all modes of operation. In hypersonic attack mode he has upgraded Cybertronian "air guardian" jet technology equipped with scramjet modules specialized for unmatched velocity. In armored ground-assault mode he transgresses all terrain and overcomes all opposition. Battle-reactive blast shields supply him with an impenetrable defense, making Optimal Optimus the most potent threat ever to face-off against Megatron and his Predacons.